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Table 1 Environmental variables used to measure hypothesized environmental drivers of body size patterns in grizzly bears within Alberta, Canada

From: Environmental, biological and anthropogenic effects on grizzly bear body size: temporal and spatial considerations

Hypothesized environmental driver and measurement variable

Units

Measurement location(s)§

Temporal scale(s)†

A. Regional habitat productivity

   

  Temperature (Winter, Spring, Summer)

°C

home range

1971-2000

  Precipitation (Winter, Spring, Summer)

mm

home range

1971-2000

  Ecosystem

categories

telemetry

C t-1

B. Inter-annual environments (deviations)

   

  Temperature (Winter, Spring, Summer)

°C

home range

Bt-1, t 0, t+1 & Ct-1, t 0

  Precipitation (Winter, Spring, Summer)

mm

home range

Bt-1, t 0, t+1 & Ct-1, t 0

C. Local habitat quality

   

  Shrub habitat (quadratic)

%

telemetry

C t-1

  Canopy cover (quadratic)

%

telemetry

C t-1

  Variation in canopy cover

%

telemetry

C t-1

  Deciduous canopy cover (quadratic)

%

telemetry

C t-1

  Forest age (quadratic)

years

telemetry

C t-1

  Forest age variation

years

telemetry

C t-1

  Regenerating forest habitat (quadratic)

%

telemetry

C t-1

  Variation in regen. forest age

years

telemetry

C t-1

  Soil wetness (quadratic)

index

telemetry

C t-1

D. Human footprint & activity

   

  Private lands

%

telemetry

C t-1

  Protected area

%

telemetry

C t-1

  Mortality risk

index

telemetry

C t-1

  Safe harbour habitat

index

telemetry

C t-1

  Linear feature density

km/km2

telemetry

C t-1

  Distance to human feature

m

telemetry

C t-1

  Distance to active energy well

m

telemetry

C t-1

E. Landscape change

   

  Annual rate of habitat change

%

telemetry

C t-1

  1. § Home ranges estimated by 50% multi-annual kernels; climate variables measured at kernel centroid; † Temporal scales relate to time of measurements; B relates to birth year & C to capture year. For inter-annual variation, 2-yrs prior to and up to 1-yr following birth or 1-yr prior to and the year of capture are considered.